Betty has a hard time telling Daniel important news about her future; Amanda copes with the death of her dog; Marc wonders if he will ever have a successful relationship.

As Hilda’s wedding day approaches, Betty asks Daniel to be her plus-one, Ignacio invites Elena, and Amanda is set on fixing up Marc with her new client, soap opera star Spencer Cannon (Bryan Batt). Meanwhile, Justin weighs whether his own plus-one could be Austin, Betty considers unexpected offers in her personal and professional life, and Wilhelmina finds a way to fight the Meade empire by using Tyler as her pawn.

Betty brings Hilda on an assignment to cover London Fashion Week; Willie joins AA to get closer to Tyler and to double-cross Claire and Daniel; Bobby wonders why Justin is spending time with Austin.

After being knocked unconscious at a photo shoot, Betty has a dream in which her orthodontist (Kathy Najimy) shows her how her life would have been had she never worn braces.

Betty is pressured into dating an annoying firefighter (Rich Sommer); Amanda persuades Claire to hire Tyler; Hilda meets Bobby’s parents.

Daniel helps Betty to cover a low-priority show. Betty discovers a new designer, Marisa, and decides to promote her however she faces stiff competition from Marc. Amanda struggles to persuade Marc or Betty to vouch for her to Wilhemina about her clothing line. Tyler tries to find Claire in New York.